Westneat, D. F. 2012. Evolution in response to social selection: The importance of interactive effects of traits on fitness. Evolution 66: 890-895.

Wetzel D.P, Westneat D. F., and I. R. K. Stewart. 2012. Heterozygosity predicts clutch and egg size but not plasticity in a wild house sparrow population with no evidence of inbreeding. Molecular Ecology 21: 406-420.

Cassone, V.M. and D. F. Westneat. 2012. The bird of time: cognition and the avian biological clock. Frontiers in Molecular Neuroscience.

Ensminger, A. L. and D. F. Westneat. 2012. Personality and plasticity in habituation and neophobia in male and female house sparrows (Passer domesticus). Ethology 118:1-11.

Lüpold, S., Birkhead, T. R. and D. F. Westneat. 2012. Seasonal variation in ejaculate traits of male red-winged blackbirds (Agelaius phoeniceus). Behavioral Ecology and Sociobiology 66: 1607-1617.



Stewart, I. R. K. and D. F. Westneat. 2013. Patterns of hatching failure in the house sparrow, Passer domesticusJournal of Avian Biology 44: 69-79.

Sousa, B. F. and D. F. Westneat. 2013. Patterns of social and extra-pair mating in a polygynous songbird, the dickcissel (Spiza americana). 

       Behavioral Ecology and Sociobiology 67: 243-255.

Westneat, D. F., Schofield, M. & Wright, J. 2013. Parental behavior exhibits between-individual variance, plasticity and heterogeneous residual variance. Behavioral Ecology 24: 598-604.

Diep, S. K. and D. F. Westneat. 2013. Integration of function and ontogeny in the evolution of status signals. Behaviour 150: 1015-1044.

Stewart, I. R. K. and D. F. Westneat. 2013. Dietary calcium, but not glutathione, affects bib size in juvenile male House Sparrows.

       The Condor,  115, 921-930.

Sousa, B. F. and Westneat, D. F. 2013. Variance in mating success does not produce strong sexual selection in a polygynous songbird. Behavioral Ecology, 24, 1381-1389.



Lüpold, S., Birkhead, T. R., and Westneat D. F. 2011. Geographical variation in sperm morphology in the red-winged blackbird (Agelaius phoeniceus). Evolutionary Ecology 25: 373-390.

Schrey, A., Grispo, M., Awad, M., Cook, M., McCoy, E., Mushinsky, H., Albayrak, T., Bensch, S., Butler, L.,  Fokidis, H., Jensen, H.,  Imboma, T., Kessler-Rios, M., Marzal, A., Stewart, I.R.K., Westerdahl, H., Westneat, D.F., Martin, L. B., Zehtindjiev, P. 2011. Broad-scale latitudinal patterns of genetic diversity among native European and introduced house sparrow (Passer domesticus) populations. Molecular Ecology 20:1133-1143.

Hartman, P.J, Wetzel, D.P., Crowley, P. H., and D. F. Westneat. 2011. The impact of extra-pair mating behavior on hybridization and genetic introgression. Theoretical Ecology 12: 219-229.

Westneat, D. F., M. I. Hatch, D. P. Wetzel, and A. L. Ensminger. 2011. Individual variation in parental care reaction norms: integration of personality and plasticity. The American Naturalist 178: 652-667. 

Stewart, I.R. K. and D. F. Westneat. 2011. Production of melanin-based plumage traits in male House Sparrows: no effect of dietary copper. International Studies of Sparrows 35: 4-15.


Stewart, S. L. M., Westneat, D. F., and Ritchison G. 2010. Extra-pair paternity in eastern bluebirds: Effects of manipulated density and natural patterns of breeding synchrony. Behavioral Ecology and Sociobiology 64:463-473. (15)

Fox, C. W. and Westneat, D. F. 2010. Adaptation. Pp. 16-31 in Westneat, D. F. and Fox, C. W. (eds) Evolutionary Behavioral Ecology, Oxford University Press.

Stewart, IRK and Westneat DF. 2010. Dietary calcium negatively affects the size of a status signal in juvenile male house sparrows (Passer domesticus). The Auk 127:549-557. 


Lynn, SG, Powell, KA, Westneat, DF, and Shepherd, BS. 2009. Seasonal and sex-specific mRNA levels of key endocrine genes in adult yellow perch (Perca flavescens) from Lake Erie. Marine Biotechnology 11: 210-222.

Dzialak, M.R., K. M. Carter, M. J. Lacki, D.F. Westneat, and K. Anderson. 2009. Activity of post-fledging peregrine falcons in different rearing and habitat conditions.  Southeastern Naturalist 8: 93-106.

Westneat DF, Stewart IRK, and Hatch MI. 2009. Complex interactions among temporal variables affect the plasticity of clutch size in a multi-brooded bird. Ecology 90:1162-1174.

Lüpold S, Linz GM, Rivers JW, Westneat DF, Birkhead TR. 2009. Sperm competition selects beyond relative testes size in birds. Evolution 63: 391-402.

Wetzel, D. P. and Westneat, D. F. 2009. Heterozygosity and extra-pair paternity: Biased tests result from the use of shared markers. Molecular Ecology 9: 2010-2021.

Westneat, DF and Sih, A. 2009. Sexual conflict as a partitioning of selection. Biology Letters 5: 675-677.

Kinnard T. B. and Westneat D. F. 2009. Phenotypic and genetic variance of house sparrows, Passer domesticus, early in development. The Auk 126: 884-895.

  2008 and earlier

Do you really want to read that old stuff? I am rather proud of some of it, so if you are curious, maybe check out my profile on Google Scholar. 


English, W. B., Schamel, D., Tracy, D. M, Westneat, D. F., and Lank, D. B. 2014. Sex ratio varies with egg investment in the red-necked phalarope (Phalaropus lobatus). Behavioral Ecology and Sociobiology 68: 1939-1949.

Liu, I. A., Johndrow, J. E., Abe, J., Lupold, S., Yasukawa, K., Westneat, D. F., and Nowicki, S. 2014. Genetic diversity does not explain variation in extra-pair paternity in multiple populations of a songbird. Journal of Evolutionary Biology 28: 1156-1169.

Wetzel, D. P. and Westneat, D. F. 2014. Parental care syndromes in house sparrows: positive covariance between provisioning and defense linked to parental identity. Ethology 120: 249-257.

Westneat, D. F., Bokony, V., Burke, T., Chastel, O., Jensen, H., Kvalnes, T., Lendvai, A. Z., Liker, A., Mock, D., Schroeder, J., Schwagmeyer, P. L., Sorci, G., Stewart, I. R. K. 2014. Multiple aspects of plasticity vary among populations of a globally distributed songbird. Journal of Animal Ecology, 83, 876-887. 


Westneat, D. F. and Fox, C. W. 2010. Evolutionary Behavioral Ecology. Oxford University Press.

​This edited book targeted for graduate students presents the foundational concepts in behavioral ecology and incorporates quantitative and molecular genetic perspectives. It includes both core hypotheses and controversies that shaped this field and describes new approaches that influence research today. The book reflects the primary role behavioral ecology has had in developing our understanding of the ecology of selection over the past 40+ years. 



Liu, I. A., Johndrow, J. E., Abe, J., Lupold, S., Yasukawa, K. Westneat, D. F., and Nowicki, S. 2015. Genetic diversity does not explain variation in extra-pair paternity in multiple populations of a songbird. Journal of Evolutionary Biology 28, 1156-1169. 

Stewart, I.R.K, Westneat, D. F., and R. L. Barrick.  2015. Food supplementation and extrapair paternity in house sparrows. American Midland Naturalist 174: 278-289.

Wetzel, D. P., Hatch, M. I. ,and D. F. Westneat. 2015. Genetic sources of individual variation in parental care behavior. Behavioral Ecology and Sociobiology 69: 1933-1943.

Westneat, D. F., Wright, J., and N. J. Dingemanse. 2015. The biology hidden inside residual within-individual phenotypic variation. Biological   Reviews, doi:10.1111/brv.12131.



Pelletier, K., Oedewaldt, C., & Westneat, D. F. (2016). Surprising flexibility in parental care revealed by experimental changes in offspring demand. Animal Behaviour, 122, 207-215.

Allegue, H., Araya-Ajoy, Y. G., Dingemanse, N. J., Dochtermann, N. A., Garamszegi, L. Z., Nakagawa, S., Réale, D., Schielzeth, H. and Westneat, D. F. (2016), Statistical Quantification of Individual Differences (SQuID): an educational and statistical tool for understanding multilevel phenotypic data in linear mixed models. Methods Ecol Evol. doi:10.1111/2041-210X.12659

Dillard J. R. and Westneat, D. F.  2016. Disentangling the correlated evolution of monogamy and cooperation. Trends in Ecology and Evolution 31: 503-513.

David F. Westneat| Tel 01 859 323 9499 | Fax 859 257 1717 | david.westneat@uky.edu


Moldoff DE and Westneat DF. 2017. Foraging sparrows exhibit individual differences but not a syndrome when responding to multiple kinds of novelty. Behavioral Ecology doi:10.1093/beheco/arx014.

Westneat DF, Mutzel A, Bonner S, and Wright J. 2017. Experimental manipulation of brood size affects several levels of phenotypic variance in offspring and parent pied flycatchers. Behavioral Ecology and Sociobiology 71:91 (DOI 10.1007/s00265-017-2315-3)

Mathot KJ, Olsen A-L, Mutzel A, Araya-Ajoy YG, Niclaus M, Westneat DF, Wright J, Kempenaers B and Dingemanse NJ. 2017. Provisioning tactics of great tits in response to long-term brood manipualtions differ across years. Behavioral Ecology.

Biagolini-Jr C, Westneat DF, and Francisco MR. 2017. Does habitat structural complexity influence the frequency of extra-pair paternity in birds? Behavioral Ecology and Sociobiology 71: 101. (DOI 10.1007/s00265-017-2329-x) 


Salzman, T., McLaughlin, A., Westneat, D. F., and P. H. Crowley. (2018) Energetic trade-offs and feedbacks between behavior and metabolism influence correlations between pace-of-life attributes. Behavioral Ecology and Sociobiology, 72: 54. doi.org/10.1007/s00265-018-2460-3.


Sánchez-Tójar, A., Nakagawa, S., Sánchez-Fortún, M., Martin, D. A., Ramani, S., Girndt, A., Bókony, V., Liker, A., Westneat, D. F., Burke, T., and J. Schroeder (2018) Meta-analysis challenges a textbook example of status signaling and demonstrates publication bias. eLife, in press. 



Westneat, D. F., Potts, L. A., Sasser, K. L, and Shaffer, J. (2019) Causes and consequences of plasticity in complex environments. Trends in Ecology and Evolution 34:555-568.

Westneat, D. F., & Mutzel, A. (2019). Variable parental responses to changes in offspring demand have implications for life history theory. Behavioral Ecology and Sociobiology, 73 (9), 130.






Araya‐Ajoy, Y.G., Westneat, D. F., and Wright, J. Pathways to social evolution and their evolutionary feedbacks. Evolution in press. 


Schielzeth, H., Dingemanse, N.J., Nakagawa, S., Westneat, D.F., Allegue, H., Teplitsky, C., Réale, D., Dochtermann, N.A., Garamszegi, L.Z. and Araya‐Ajoy, Y.G., Robustness of linear mixed‐effects models to violations of distributional assumptions. Methods in Ecology and Evolution, online.