Stieha C.,  A. Middleton, J. Stieha; S. Trott, D. N. McLetchie. 2014. The dispersal process of asexual propagules and the contribution to population persistence in Marchantia (Marchantiaceae). American Journal of Botany 101:348-356; doi:10.3732/ajb.1300339

Brzyski, J. R., W. Taylor, D. N. McLetchie, 2014, Reproductive allocation between the sexes, across natural and novel habitats, and its impact on genetic diversity. Evolutionary Ecology 28:247-261


Brzyski, J., K. Adams, C. Walter, K. H. Gale, D. N. McLetchie 2012. Characterization of 12 polymorphic microsatellite markers in the liverwort Marchantia inflexa (Marchantiaceae) American Journal of Botany (vol. 99 no. 11 e440-e442). http://www.amjbot.org/cgi/doi/10.3732/ajb.1200187

Stark L. R., J C Brinda, D. N. McLetchie, M J Oliver. 2012. Extended periods of hydration do not elicit dehardening to desiccation tolerance in regeneration trials of the moss Syntrichia caninervis. International Journal of Plant Sciences 173 333-343. 10.1086/663970


Reynolds, L. A., and D N. McLetchie. 2011. Short distance between extreme microhabitats do not result in ecotypes in Syntrichia caninervis, Journal of Bryology 33: 148-153 10.1179/1743282011Y.0000000004

Benassi, M. L., L. R. Stark, J. C. Brinda, D. N. McLetchie, M. Bonine, and B.D. Mishler. 2011. Plant size, sex expression, and sexual reproduction along an elevation gradient in a desert moss. The Bryologist 114:277-288 10.1639/0007-2745-114.2.277

Horsley K., L. R. Stark, and D. N. McLetchie. 2011. Does the silver moss Bryum argenteum exhibit sex-specific patterns in vegetative growth rate, asexual fitness, or prezygotic reproductive investment? Annals of Botany 107: 897-907 doi: 10.1093/aob/mcr027

Stark L. R., J C Brinda, D. N. McLetchie. 2011.Effects of increased summer precipitation and N deposition on Mojave Desert populations of the biological crust moss Syntrichia caninervis, Journal of Arid Environments 75: 457-463  doi:10.1016/j.jaridenv.2010.12.015


Stark, L. R., D. N. McLetchie, S. D. Smith and M. J. Oliver. 2011. Responses of a biological crust moss to increased monsoon precipitation and N deposition in the Mojave Desert. pp. 149-168 in Tuba Z, Slack N, and Stark L. (eds), Bryophyte Ecology and Climate Change. Cambridge University Press. Cambridge. (Book Chapter)




Stark, L. R., D. N. McLetchie, and S. M. Eppley. 2010. Sex ratios and the shy male hypothesis in the moss Bryum argenteum (Bryaceae) The Bryologist 113: 788-797 (http://www.bioone.org/doi/full/10.1639/0007-2745-113.4.788)

Groen, K. E., Stieha, C., Crowley, P. H., and McLetchie, D. N. 2010. Sex-specific plant responses to light intensity and canopy openness: implications for spatial segregation of the sexes. Oecologia 162: 561-570 doi:10.1007/s00442-009-1473-z

Groen, K. E., Stieha, C., Crowley, P. H., and McLetchie, D. N. 2010. Sex-specific plant responses to two light levels in the liverwort Marchantia inflexa (Marchantiaceae). The Bryologist 113: 81-89.

Cole, C., Stark, L. R., Bonine M., and McLetchie, D. N. 2010. Transplant survivorship of bryophyte soil crusts in the Mojave Desert. Restoration Ecology 18: 198-205. doi:10.1111/j.1526-100X.2008.00445.x.




Stark, L. R, Brinda, J. and McLetchie, D. N. 2009. An experimental demonstration of the cost of sex and a potential resource limitation on reproduction in the moss Pterygoneurum (Pottiaceae). American Journal of Botany 96: 1-11. doi:10.3732/ajb.0900084

Stark, L. R, McLetchie D. N. and Roberts S. 2009. Gender differences and a new adult eukaryotic record for upper thermal tolerance in the desert moss Syntrichia caninervis Journal of Thermal Biology 34: 131-137.  doi:10.1016/j.jtherbio.2008.12.001




Moore J.D., L. Kollar, D.N. McLetchie. 2016. Does selection for gamete dispersal and capture lead to a sex difference in clump water-holding capacity? American Journal of Botany. 103: 1449-1457 doi:10.3732/ajb.1600096.


Rose M., J Burton D.N. McLetchie. 2016. Sex differences and plasticity in dehydration tolerance: insight from a tropical liverwort. Annals of Botany doi: 10.1093/aob/mcw102.

Stieha C., G García-Ramos, D.N. McLetchie, P. Crowley. 2016. Maintenance of the Sexes and Persistence of a Clonal Organism in Spatially Complex Metapopulations.  Evolutionary Ecology DOI 10.1007/s10682-016-9841-8.


Stark L. R, D. N. McLetchie, S. M. Eppley and J. L. Greenwood. 2016. Moss Antheridia are Desiccation Tolerant: Rehydration Dynamics Influence Sperm Release in Bryum argenteum (Bryaceae) American Journal of Botany doi:10.3732/ajb.1600026


Publications 2009 to present - For earlier pubs see Google scholar


Designed by P

For earlier pubs see Google scholar





D. Nicholas McLetchie Ph.D., Phone: 859-257-6786, Fax: 859-257-1717,  Email: mclet@uky.edu
Department of Biology, 101 Morgan Building, University of Kentucky, Lexington KY 40502-0225


DN McLetchie, G García-Ramos. 2017. A Predictive Relationship between Population and Genetic Sex Ratios in Clonal Species, Acta Oecologica 80, 18-23.